In yeast, LYS5, ADE1, and URA2 are genes required to synthesize lysine, adenine and uracil, respectively. A MATa lys5 ADE1 URA2 haploid was mated to a MATA LYS5 ade1 ura2 haploid to make a triply heterozygous diploid. This diploid was put through meiosis, 100 tetrads were dissected and the spore colonies transferred to medium lacking either lysine, adenine or uracil to determine which spores were prototrophic or auxotrophic for those nutrients. The following results were obtained: LYS5-ADE1 ADE1-URA2 LYS5-URA2 PD 20 18 50 NPD I 58 62 42 22 20 8
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- A heterozygous diploid yeast Aa Bb went through meiosis. What percentage of the haploid spores will have recombinant combinations of alleles? What if genes A and B are unlinked? Explain What is genes A and B are linked? ExplainMeiosis is characterized by the pairing of homologouschromosomes during prophase I. In many species, an elaboratestructure called the synaptonemal complex forms betweenhomologues. During this pairing, homologues may exchangechromosomal material at sites called chiasmata. In meiosis I, thehomologues separate from each other, reducing the chromosomenumber to the haploid state (thus the reductive division). It isfollowed by a second division without replication, during whichsister chromatids become separated. The result of meiosis I and IIis four haploid cells. If sister chromatids separated at the first division, would meiosis still work?Map distances were determined for four differentgenes (MAT, HIS4, THR4, and LEU2) on chromosome III of the yeast Saccharomyces cerevisiae:HIS4 ↔ MAT 37 cMTHR4 ↔ LEU2 35 cMLEU2 ↔ HIS4 23 cMMAT ↔ LEU2 16 cMMAT ↔ THR4 20 cMWhat is the order of genes on the chromosome?
- For a haploid fungus, the starting point in the biosynthesis of the amino acid arginine is Compound X, which is always present in and absorbed from the environment. The arginine biosynthetic pathway is: Enzyme A Enzyme B Enzyme Ç Compound X It is know that genes encoding enzymes A and C are on two different chromosomes. Compound Y Compound Z- Arginine A mutant strain of genotype a (lacking only enzyme A) is crossed to a mutant strain of genotype c (lacking only enzyme C) to generate a diploid strain. Sporulation (i.e. meiosis) is subsequently induced in the resulting diploid strain. What proportion of the spores (haploids formed by sporulation) is expected to grow on medium without arginine but supplemented with Compound Y? O 100% 50% 0% 25%One reason that worker bees forgo their own reproduction to help their sister (the queen) reproduce is that female bees are more closely related to their sisters than they are to their own offspring. This quirk of genetics results from the fact that bees have haplodiploid sex determination, in which females are diploid, with a mother and a father, but males are haploid, developing from unfertilized eggs. Because males are haploid, they produce sperm by mitosis. Explainwhy haplodiploid sex determination causes females to be more closely related to their sisters than to their offspring.An individual heterozygous for a reciprocal translocation possesses the following chromosomes: A B • C D E F G A B • C D V W X R S • T U E F G R S • T U V W X Q. Draw the pairing arrangement of these chromosomes in prophase I of meiosis.
- Shown below are photomicrographs of Rhoeo tradescantia cells undergoing meiosis. Answer the following question for each of the photomicrographs: Identify the cytogenetic abnormality observed (ex. ring, chain, laggard, bridge). Identify the meiotic stage in which these aberrations are observed (as shown in the photomicrograph). Explain how these aberrations are formed and relate to the possible causal mutation(s). Will this result to sterile and/or fertile gametes? Explain.In the haploid yeast Saccharomyces cerevisiae, the twomating types are known as MATa and MATα. You cross apurple (ad-) strain of mating type a and a white (ad+)strain of mating type α. If ad- and ad+ are alleles of onegene, and a and α are alleles of an independently inherited gene on a separate chromosome pair, what progenydo you expect to obtain? In what proportions?You cross two yeast strains one is an ade auxotroph the other is a pro auxotroph and allow the diploid to sporulate. When you score each spore in the ascus you find the following proportions: 518 PD, 8 NPD, and 225 T. a.) What are the genotypes of each spore in all three types of the tetrads. b) Are these genes linked why or why not? c.) If these genes are unlinked what would you expect the progeny numbers and ratios to be? d.) What is the formula to determine the most accurate distance between these genes? If linked what is the map distance?
- In Neurospora, his2 mutants require the amino acidhistidine for growth, and lys4 mutants require theamino acid lysine. The two genes are on the samearm of the same chromosome, in the ordercentromere - his2 - lys4.A his2 mutant is mated with a lys4 mutant. Draw all ofthe possible ordered asci that could result from meioses in which the following events occurred, accountingfor the nutritional requirements for each ascospore.Ascospores without any copy of a chromosome willabort and die, turning white in the process.a. A single crossover between the centromere and his2b. A single crossover between his2 and lys4c. Nondisjunction during the first meiotic divisiond. Nondisjunction during the second meiotic divisione. A single crossover between the centromere andhis2, followed by nondisjunction during the firstmeiotic divisionf. A single crossover between his2 and lys4,followed by nondisjunction during the firstmeiotic divisionDuring metaphase I of meiosis, tetrads align along the metaphase plate independently of each other. Therefore, there is a random “shuffle” of maternal and paternal chromosomes in the resulting gametes.The following diagram demonstrates how this works in a diploid cell with four chromosomes (2n=4) . Because there are two pairs of chromosomes and each pair can align in one of two ways during metaphase I, the number of possible variations in the gametes produced is 22 or 4.For an organism that is , there are three pairs of chromosomes, so the number of possible variations in the gametes produced due to independent assortment in metaphase I is 23 or 8. In an organism with a haploid number of 7, how many possible combinations of maternal and paternal chromosomes can occur in its gametes? a. 72=49 b. 27=128 c.17=1 d. 214=16 384Consider the following two meiocytes in metaphase I, with crossover positions as indicated for cell A. Solid black lines indicate spindle microtubules. Assume blue chromosomes represents paternal chromosomes and green maternal chromosomes. F G N R A B D e g a B d E Cell A Cell B i) How many chromosomes and chromatids were present in cell A during the G1 phase? Please clearly specify chromosomes and chromatids in your answer. ii) For cell A, list all the gametes that will be produced from this cell. Give the alleles of each gamete contained within a set of brackets. You may ignore independent assortment. iii) For cell B, list all the gametes that will be produced from this cell given independent assortment. Give the alleles of each gamete contained within a set of brackets. You may assume that no crossing over would take place.